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75 Although ACP transfer catalyzed by Tsr3 is the final step in m 1 acp 3 Ψ modification during 18S rRNA maturation, studies of yeast mutants lacking enzymes for pseudouridylation and methylation have suggested that in vivo ACP transfer activity is independent of either modification such that the corresponding mutants can produce acp 3 U (27) or acp 3 Ψ, respectively